Peer-Reviewed Articles (Click link for PDF)


41) Van Wittenberghe S, Alonso L, Malenovský Z, Moreno J. In vivo photoprotection mechanisms observed from leaf spectral absorbance changes showing VIS–NIR slow-induced conformational pigment bed changes. Photosynthesis Research 142:3. DOI: 10.1007/s11120-019-00664-3 (PDF: Van Wittenberghe et al 2019)

40) Bacour C, Maignan F, MacBean N, Porcar‐Castell A, Flexas J, Frankenberg C, Peylin P, Chevallier F, Vuichard N, Bastrikov V. (2019) Improving estimates of Gross Primary Productivity by assimilating solar‐induced fluorescence satellite retrievals in a terrestrial biosphere model using a process‐based SIF model. Journal of Geophysical Research: Biogeosciences. DOI : 10.1029/2019JG005040 (PDF: Bacour et al 2019)

39) Liu W, Atherton J, Mõttus M, Gastellu-Etchegorry JP, Malenovský Z, Raumonen P, Åkerblom M, Mäkipää R, Porcar-Castell A. (2019) Simulating solar-induced chlorophyll fluorescence in a boreal forest stand reconstructed from terrestrial laser scanning measurements. Remote Sensing of Environment. DOI: 10.1016/j.rse.2019.111274 (PDF: Liu et al 2019)

38) Atherton J, Liu W, Porcar-Castell A. (2019) Nocturnal Light Emitting Diode Induced Fluorescence (LEDIF): A new technique to measure the chlorophyll a fluorescence emission spectral distribution of plant canopies in situ. Remote Sensing of Environment. DOI: 10.1016/j.rse.2019.03.030 (PDF: Atherton et al 2019)

37) Féret JB, Le Maire G, Jay S, Berveiller D, Bendoula R, Hmimina G, Cheraiet A, Oliveira JC, Ponzoni FJ, Solanki T, de Boissieu F, Chave J, Nouvellon Y, Porcar‐Castell A, Proisy C. Soudani K, Gastellu-Etchegorry JP, Lefèvre-Fonollosa, MJ. (2019) Estimating leaf mass per area and equivalent water thickness based on leaf optical properties: Potential and limitations of physical modeling and machine learning. Remote Sensing of Environment. DOI: 10.1016/j.rse.2018.11.002 (PDF: Féret et al 2019)

36) Zhang C, Atherton J, Peñuelas J, Filella I, Kolari P, Aalto J, Ruhanen H, Bäck J, Porcar‐Castell A. (2019) Do all chlorophyll fluorescence emission wavelengths capture the spring recovery of photosynthesis in boreal evergreen foliage?. Plant, cell & environment. DOI: 10.1111/pce.13620  (PDF: Zhang et al 2019)

35) Raczka B, Porcar‐Castell A, Magney T, Lee JE, Köhler P, Frankenberg C, Grossmann K, Logan BA, Stutz J, Blanken PD, Burns SP, Duarte H, Yang X, Lin JC, Bowlin DR. (2018) Sustained Non‐Photochemical Quenching Shapes the Seasonal Pattern of Solar‐Induced Fluorescence at a High‐Elevation Evergreen Forest. Journal of Geophysical Research: Biogeosciences. DOI: 10.1029/2018JG004883 (PDF: Raczka et al 2019)

34) Magney TS, Frankenberg C, Köhler P, North G, Davis TS, Dold C, Dutta D, Fisher JB, Grossmann K, Harrington A, Hatfield J, Stutz J, Sun Y, Porcar-Castell A. (2019) Disentangling changes in the spectral shape of chlorophyll fluorescence: Implications for remote sensing of photosynthesis. Journal of Geophysical Research: Biogeosciences. DOI: 10.1029/2019JG005029 (PDF: Magney et al 2019)

33) Rajewicz PA, Atherton J, Alonso L, & Porcar-Castell A. (2019). Leaf-level spectral fluorescence measurements: comparing methodologies for broadleaves and needles. Remote Sensing, 11(5), 532. DOI: 10.3390/rs11050532 (PDF: Rajewicz et al 2019)

32) Nichol CJ, Drolet G, Porcar-Castell A, Wade T, Sabater N, Middleton EM, MacLellan C, Levula J, Mammarella I, Vesala T, & Atherton J. (2019). Diurnal and Seasonal Solar Induced Chlorophyll Fluorescence and Photosynthesis in a Boreal Scots Pine Canopy. Remote Sensing, 11(3), 273. DOI: 10.3390/rs11030273 (PDF: Nichol et al 2019)

31) Solanki T, Aphalo P J, Neimane S, Hartikainen SM, Pieristè M, Shapiguzov A, Porcar-Castell A, Atherton J, Heikkilä A, & Robson TM. (2019). UV-screening and springtime recovery of photosynthetic capacity in leaves of Vaccinium vitis-idaea above and below the snow pack. Plant physiology and biochemistry, 134, 40-52. DOI: 10.1016/j.plaphy.2018.09.003 (PDF: Solanki et al 2019 )


30) Sabater N, Vicent J, Alonso L, Verrelst J, Middleton E, Porcar-Castell A, & Moreno J (2018). Compensation of oxygen transmittance effects for proximal sensing retrieval of canopy–leaving sun–induced chlorophyll fluorescence. Remote Sensing, 10(10), 1551.DOI: 10.3390/rs10101551 (PDF: Sabater et al 2018 )

29) Fernández-Marín B, Atherton J, Olascoaga B, Kolari P, Porcar-Castell A, & García-Plazaola, JI (2018). When the sun never sets: daily changes in pigment composition in three subarctic woody plants during the summer solstice. Trees, 32(2), 615-630. DOI: 10.1007/s00468-018-1660-9 (PDF: Fernández-Marín et al 2018 )


28) Sun Y, Frankenberg C, Wood JD, Schimel DS, Jung M, Guanter L, Drewry DT, Verma M, Porcar-Castell A, Griffins TJ, Gu L, Magney TS, Köhler P, Evans B, Yuen K (2017) OCO-2 advances photosynthesis observation from space via solar-induced chlorophyll fluorescence. Science 358. Doi: 10.1126/science.aam5747 (PDF: Sun et al. 2017)

27) Jokinen T, Kontkanen J, Lehtipalo K, Manninen HE, Aalto J, Porcar-Castell A,.. and 17 more (2017) Solar eclipse demonstrating the importance of photochemistry in new particle formation. Scientific Reports, DOI: 10.1038/srep45707 (PDF: Jokinen et al. 2017)

26) Atherton J, Olascoaga B, Alonso L, Porcar-Castell A (2017) Spatial variation of leaf optical properties in a boreal forest is influenced by species and light environment. Frontiers in Plant Science, doi: 10.3389/fpls.2017.00309 (PDF: Atherton et al. 2017)

25) Ulsig L, Nichol CJ, Huemmrich KF, Landis DR, Middleton EM, Lyapustin AI, Mammarella I, Levula J, Porcar-Castell A (2017) Detecting inter-annual variations in the phenology of evergreen conifers using long-term MODIS vegetation index time series. Remote Sensing, doi: 10.3390/rs9010049 (PDF: Ulsig et al. 2017)


24) Olascoaga B, Mac Arthur A, Atherton J, Porcar-Castell A (2016). A comparison of methods to estimate photosynthetic light absorption in leaves with contrasting morphology. Tree Physiology doi:10.1093/treephys/tpv133 (PDF: Olascoaga et al. 2016)

23) Atherton J, Nichol C, Porcar-Castell A (2016). Using Spectral Chlorophyll Fluorescence and the Photochemical Reflectance Index to predict Physiological Dynamics. Remote Sensing of Environment 176: 17-30. (PDF: Atherton et al. 2016)


22) Lindfors L, Hölttä T, Lintunen A, Porcar-Castell A, Nikinmaa E, Juurola E (2015). Dynamics of leaf gas exchange, chlorophyll fluorescence and stem diameter changes during freezing and thawing of Scots pine seedlings. Tree Physiology. (PDF: Lindfors et al. 2015)

21) Porcar-Castell A, MacArthur A, Rossini M, Eklundh L, Pacheco-Labrador J, Anderson K, Balzarolo M, Martín MP, Jin H, Tomelleri E, Cerasoli S, Sakowska K, Hueni A, Julitta T, Nichol CJ, Vescovo L. EUROSPEC: At the interface between Remote Sensing and CO2 flux measurements in Europe. Biogeosciences 12: 6103-6124. (PDF: Porcar-Castell et al. 2015)

20) Wingate L, Ogée J, Cremonese E, Filippa G, Mizunuma T, Migliavacca M, Moisy C, Wilkinson M, Moureaux C,Wohlfahrt G, Hammerle A, Gimeno G, Porcar-Castell A, Galvagno M, Nakaji T, Morison J, Kolle O, Knohl A, Kutsch W, Kolari P, Nikinmaa E, Ibrom A, Gielen B, Eugster W, Balzarolo M, Papale D, Klumpp K, Köstner B, Grünwald T, Joffre R, Ourcival J-M, Hellstrom M, Charles G, Longdoz B, Genty B, Levula J, Heinesch B, Sprintsin M, Yakir D, Manise T, Guyon D, Ahrends H, Plaza-Aguilar A, Hong Guan J, Grace J (2015). Interpreting canopy development and physiology using the EUROPhen camera network at flux sites. Biogeosciences Discussions 12: 7979-8034. (PDF: Wingate et al. 2015)

19) Cendrero-Mateo MP, Carmo-Silva AE, Porcar-Castell A, Nearing GS, Hamerlynck EP, Papuga SA, Moran MS (2015). Dynamic response of plant chlorophyll fluorescence to light, water and nutrient availability. Functional Plant Biology 42:746-757 (PDF:Cendrero-Mateo et al. 2015)

18) Aalto J, Porcar-Castell A, Atherton J, Kolari P, Pohja T, Hari P, Nikinmaa E, Petäjä T, Bäck J (In Press). Onset of photosynthesis in spring speeds up monoterpene synthesis and leads to emission bursts. Plant, Cell and Environment. doi: 10.1111/pce.12550 (PDF: Aalto et al. 2015)


17) Kolari P, Chan T, Porcar-Castell A, Bäck J, Nikinmaa E, Juurola E, 2014. Field and controlled environment measurements show strong seasonal acclimation in photosynthesis and respiration potential in boreal Scots pine. Frontiers in Plant Science, doi: 10.3389/fpls.2014.00717 (PDF: Kolari et al. 2014)

16) Porcar-Castell A, Tyystjärvi E, Atherton J, van der Tol C, Pfündel E, Flexas J, Frankenberg C, Moreno J, Berry JA, 2014. Linking chlorophyll fluorescence to photosynthesis for remote sensing applications: mechanisms and challenges (Darwin Review). Journal of Experimental Botany 65: 4065-4095. (PDF:Porcar-Castell et al. 2014 and the corrigendum Porcar-CAstell et al. 2015b )

15) Olascoaga B, Juurola E, Pinho P, Nikinmaa E, Halonen L, Bäck J, Porcar-Castell A, 2014. Seasonal variation in the reflectance of photosynthetically active radiation from epicuticular waxes of Scots pine (Pinus sylvestris L.) needles. Boreal Environment Research 19(B):132-141. (PDF: Olascoaga et al. 2014)

14) Drolet G, Wade T, Nichol CJ, MacLellan C, Levula J, Porcar-Castell A, Nikinmaa E, Vesala T, 2014. A low-cost temperature controlled spectrometer system for continuous and unattended measurements of canopy spectral radiance and reflectance. International Journal of Remote Sensing 35:1769-1785.(PDF:Drolet et al. 2014 )


13) Porcar-Castell A, Palmroth S, 2012. Modelling photosynthesis in highly dynamic environments: The case of sunflecks. Tree Physiology 32: 1062-1065. (PDF:Porcar-Castell & Palmroth 2012 )

12) Kuusinen N, Kolari P, Levula J, Porcar-Castell A, Stenberg P, Berninger F. 2012. Seasonal variation in boreal pine forest albedo. Agricultural and Forest Meteorology 164: 53-60. (PDF: Kuusinen et al. 2012)

11) Garcia-Plazaola JI, Esteban R, Fernández-Marín E, Kranner I, Porcar-Castell A, 2012. Thermal energy dissipation and xanthophyll cycles, beyond the Arabidopsis model. Photosynthesis Research 113:89-103. (PDF: Garcia-Plazaola et al. 2012)

10) Porcar-Castell A, García-Plazaola JI, Nichol C, Kolari P, Olascoaga B, Kuusinen N, Fernández-Marin B, Minna Pulkkinen, Juurola E, Nikinmaa E, 2012. Physiology of the seasonal relationship between photochemical reflectance index and photosynthetic light use efficiency. Oecologia 170:313-323. (PDF: Porcar-Castell et al. 2012)

9) Hölttä T, Juurola E, Lindfors L, Porcar-Castell A, 2012. Cavitation induced by a surfactant leads to a transient release of water stress and subsequent run away embolism in Scots pine (Pinus sylvestris) seedlings. Journal of Experimental Botany 63:1057-1067. (PDF:Holttä et al. 2011 )


8) Porcar-Castell A, 2011. A high-resolution portray of the annual dynamics in photochemical and non-photochemical quenching in needles of Pinus sylvestris. Physiologia Plantarum 143: 139-153. (PDF: Porcar-Castell 2011


7) Porcar-Castell A, Peñuelas J, Owen S, Llusià J, Munné-Bosch S, Bäck J, 2009. Leaf carotenoid concentrations and monoterpene emission capacity under acclimation of the light reactions of photosynthesis. Boreal Environment Research 14:794-806. (PDF: Porcar-Castell et al. 2009  Note typographic errors in equations, see “Errata… Sorry”)

6) Filella I, Porcar-Castell A, Munné-Bosch S, Bäck J, Garbulsky M, Peñuelas J, 2009. PRI assessment of long-term changes in carotenoids/chlorophyll ratio and short-term changes in de-epoxidation state of the xanthophyll cycle. International Journal of Remote Sensing 30: 4443-4455. (PDF: Filella et al. 2009)


5) Porcar-Castell A, Juurola E, Ensminger I, Berninger F, Hari P, Nikinmaa E, 2008. Seasonal acclimation of photosystem II in Pinus sylvestris. II. Studying the effect of light environment through the rate constants of sustained thermal dissipation and photochemistry. Tree physiology 28:1483-1491. (PDF: Porcar-Castell et al. 2008c)

4) Porcar-Castell A, Juurola E, Nikinmaa E, Berninger F, Ensminger I, Hari P, 2008. Seasonal acclimation of photosystem II in Pinus sylvestris. I. Estimating the rate constants of sustained thermal dissipation and photochemistry. Tree physiology 28:1475-1482. (PDF: Porcar-Castell et al. 2008b)

3) Porcar-Castell A, Pfündel E, Korhonen JFJ, Juurola E, 2008.A new monitoring PAM fluorometer (MONI-PAM) to study the short- and long-term acclimation of photosystem II in field conditions. Photosynthesis Research 96:173-179. (PDF: Porcar-Castell et al. 2008a)


2) Metslaid M, Jõgiste K, Nikinmaa E, Moser K, Porcar-Castell A, 2007. Tree variables related to growth response and acclimation of advance regeneration of Norway spruce and other coniferous species after release. Forest Ecology and Management 250:56-63. (PDF: Metslaid et al. 2007)


1) Porcar-Castell A, Bäck J, Juurola E, Hari P, 2006. Dynamics of the energy flow through PSII under changing light environments: a model approach. Functional Plant Biology 33(3): 229-239. (PDF: Porcar-Castell et al. 2006)


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